Author: (Pallas, 1811)

Zosterisessor ophiocephalus (Pallas, 1811)

Diagnosis: see genus. Dl VI (V-VII); D2 1 + 14-15 (13-16); A I + 14-15 (12-16); P 18-19 (17-20). Scales in lateral series 59-64 (53-68). Vertebrae 28 (27-29). Colour: greenish-yellow to olive, body with 12-14 irregular brownish vertical bands down sides; small dark spot on lateral midline at base of caudal fin and in upper corner of pectoral fin lobe. Size: to 24.5 cm.

Habitat: inshore, brackish water of estuaries and lagoons, in and near holes among grass roots on mud substrates. Food: small fish (sandsmelt, gobies) and larger crustaceans (such as Leander). Reproduction: April-May (Tuzlov estuaries, Black Sea) or July (Varna), spawning twice. Eggs 2.6 x 0.8 mm, in weed; fecundity 1O,OO0-45,000. Sexually mature at 2 or 3 years. Lifespan: 5 years.

Distribution: Mediterranean, Black Sea and Sea of Azov.

Eggs, larvae and young stages. No data.
Otoliths (sagitta). No data.

Kingdom Animalia
Phylum Chordata
Subphylum Vertebrata
Superclass Gnathostomata
Class Osteichthyes
Order Perciformes
Suborder Gobioidei
Family Gobiidae
Genus Zosterisessor
Species Zosterisessor ophiocephalus

Status in World Register of Marine Species

Accepted name: Zosterisessor ophiocephalus (Pallas, 1814)

Scientific synonyms and common names

Zosterisessor ophiocephalus (Pallas, 1811)

Synonyms:

Gobius ophiocephalus Pallas, 1811, Zoogr. rosso.-asiat., 3: 153 (coasts of Crimea).
? Gobius viridis Otto, 1821, Consp. anim., (1): 7; Isis (Oken), 2: [7-12] ('Habitat in Mari circa Niceam').
? Gobius filamentosus Risso, 1826, Hist. nat. Eur. mérid., 3: 284 (Nice) (see Moreau, 1 892: 1 69).
Gobius reticulatus Eichwald, 1831, Zool. spec., 3: 77 (Odessa).
Gobius lota Valenciennes, 1837, in Cuv. Val., Hist. nat. Poiss., 12: 27 (Bologna, Italy). Gobiusgous Nardo (ex Chieregini MS), 1847, Sin. mod. Spec. Lag. Ven.: 119 (Venice). Gobius venetiarum Nardo, 1860, Prosp. sist. Anim. Prov. Ven. Mar. Adr. (Venice). Gobius lota: Moreau, 1881: 200, fig. 101(7.); 1892: 167.
Gobius filamentosus: Moreau, 1892: 169.
Gobius ophiocephalus: Carus, 1893: 680 Ninni, 1938: 149, 169, fig. 42-43, 48-49, pl. XVIIIa-f Berg, 1949: 1080; Cavinato, 1952: 40, 47, 51, fig. 2D, 11 Soljan, 1963: 219, 385, 399 Zei, 1963: 550, pl. 204 Georghiev, 1966: 181, fig. 19-21
Zambriborsch, 1968: 42.
Gobius ophiocephalus forma ophiocephalus: Smitt, 1900: 548.
Gobius cephalarges: Antipa, 1909: pl. III (fig. 22a-22b) (nec Pallas, 1811).
Zostericola ophiocephalus: lljin, 1927c: 142.
Gobius (Zostericola) ophiocephalus: de Buen, 1930a: 139; 1931: 52, 68 Borcea, 1934: 174, fig. 66-70, pl. IV (fig. 23), pl. V (fig. 28, 31).
Zosterissessor ophiocephalus: Whitley, 1935: 250 Iljin, 1956: 192 Bini, 1969, 7: 125. Gobius ophiocephalus var. citrina Ninni, 1938: 154 (Sea of Marmora).
Neogobius cephalarges: Berg, 1949: fig. 818.
Gobius veneziarum Cavinato, 1952, Arch. Oceanogr. Limnol., 7 (2-3): 51 (error for G. venetiarum Nardo, 1860).
Gobius ophicephalus Iljin, 1956, Vopr. Ikhtiol., 7: 186 (error for G. ophiocephalus Pallas, 181 1).
Gobius (Zosterisessor) ophiocephalus: Banarescu, 1964: 830, fig. 367 Svetovidov, 1964: 427, 454, fig. 165.


Cammon names:

bychok-travjanik [Ru]
ghiozzo gò [It]
glavoc travas [Hr]
gobie lote [Fr]
grass goby [En]
guvid de iarba [Ro]
trevno popce [Bu]

Antipa, G. 1909. Fauna ichtiologica a Romaniei, Acad. Romand, Publ Fond. Vasili Adamachi, 16: ix+294 p., pl. I-XXXI (fig. 1-149).

Banarescu, P. 1964. Pisces. Osteichthyes. Fauna Repub. pop. rom., Bucuresti, 13: 1-962, 402 fig.

Berg, L. S. 1948-1949. Ryby presnykh vod SSSR i sopredel'nykh stran, 4 ed., 1, Opred. pofaune SSSR, no. 27, 1948: pp. 1-466, fig. 1-281; 2, ibid., no. 29, 1949: pp. 467-925, fig. 282-674; 3, ibid., no. 30, 1949: pp. 927-1382, fig. 675-946, map (in Russian).

Bini, G. 1967-1972. Atlante dei pesci delle Coste italiane. Mondo Sommerso, Milano, 9 vol: I, 1967, Leptocardi, Ciclostomi, Selaci, 206 pp., 66 fig. + 64 col. fig. II, 1971, Osteitti (Acipenseriformi, Clupeiformi, Mictofiformi, Anguilliformi), 300 pp., 73 col. fig. III, 1970, Notacantiformi . . . Zeiformi, 229 pp., 34 fig. + 63 col. fig. IV, 1968, Perciformi (Mugiloidei, Percoidei), 163 pp., 34 fig. + 49 col. fig. V, 1968, Perciformi (Percoidei), 175 pp., 22 fig. + 56 col. fig. VI, 1968, Perciformi (Trichiuroidei . . . Blennioidei), 177 pp., 48 fig + 57 col. fig. VII,1969, Perciformi (Ofidioidei . . . Dactilopteroidei), 196 pp., 57 fig. + 59 col. fig. VIII,1968, Pleuronettiformi, Echeniformi, Gobioesociformi, Tetraodontiformi, Lofiformi, 164 pp., 34 + 63 fig. IX, 1972, Introduzione. Parte generale. Aggiornamenti. Indici. 176 p.

Borcea, I. 1934a. Révision systématique et distribution géographique des Gobiidés de la mer Noire et particulièrement des eaux roumaines. Annls. scient. Univ. Jassy, 19 : 231 p., 91 fig.

Buen, F. de 1930a. Sur une collection de Gobiinae provenant du Maroc. Essai de synopsis des espèces de l'Europe. Bull. Soc. Sci. nat. phys., Maroc, 10: pp. 120-147.

Carus, J. V. 1893. Vertebrata. 1. class. Pisces: pp. 498-711, In: Prodromus faunae Mediterraneae sive Descriptio Animalium Maris Mediterranei incolarum quam comparata silva rerum quatenus innotuit adiectis locis et nominibus vulgaribus. . . Stuttgart, 1889-1893, 2: ix+854 pp.

Cavinato, G. 1952. Revisione dei Gobius della Laguna Veneta. Archo Oceanogr. LimnoL, Roma., 7 (2-3): 56 p

Cuvier, G.; Valenciennes, A. 1828-1849. Histoire naturelle des Poissons, Paris-Strasbourg, 22 vol., 11030 pp., 621 pl. (numb. 1-650).1,1828: xvi+573 pp., pl. I-VIII [author: Cuvier]; 2, 1828: xxi+490 pp., pl. 9-40 [Cuv.: pp. 1-238, pp. 249-262, pp. 387-490; Val.: pp. 238-249, pp. 262-386]; 3 April 1829: xxiii+500 pp., pl. 41-71 [Cuvier]; 4, 1829: xxvi+518 pp., pl. 72-99 [Cuvier]; 5, 1830: xxviii+499 pp., pl. 100-140 [Cuvier]; 6, 1830: xxiv+pp.559 pp., pl. 141-169 [Val.: pp. 1-425, 493-559; Cuv.: 426-491]; 7, 1831: xxix + 531 pp., pl. 170-208 [Cuv.: 1-440; Val.: pp. 441-531]; 8, 1832: xix+509 pp., pl. 209-245 [Cuv.: pp. 1-470; Val.: pp. 471-509]; 9, 1833: xxix+512 pp., pl. 246-279 [Cuv.: pp. 1-198, pp. 330-359, pp. 372-427; Val.: pp. 199-329, pp. 359-371, pp. 429-512]; 10, 1835: xxiv+482 pp., pl. 280-306 [Val.]; 11, 1836: xx+506 pp., pl. 307-343 [Val.]; 12, 1837: xxix+507 pp., pl. 344-368 [Val.]; 13, 1839: xix+505 pp., pl. 369-388 [Val.]; 14, 1840: xxii+464 pp., pl. 389-420 [Val.]; 15, 1840: xxi + 540 pp., pl. 421-455 [Val.]; 16, 1842: xx+472 pp., pl. 456-487 [Val.]; 17, 1844: xxiii+497 pp., pl. 487-519 [Val.]; 18, 1846: xix+505 pp., pl. 520-553 [Val.]; 19, 1847: xix+544 pp., pl. 554-590 [Val.]; 20, 1847: xviii+472 pp., pl. 591-606 [Val.]; 21, 1848: xiv+536 pp., pl. 607-633 [Val.]; 22, 1849: xx+532 pp., pl. 634-650 [Val.].

Eichwald, C. E. von 1831. Zoologia specialis quam expositis animalibus tum vivis . . . adidit D. Eduardus Eichwald. Pars posterior: 404 pp., 1 pl. + l pl. n. num. p. 342 [Pisces: 10-116].

Georghiev, J. M. 1966. Composition d'espèce et caractéristique des Gobiides (Pisces) en Bulgaria. Izv. nauchno-issled. Inst. Rib. Stop. Okeanogr. Varna, 7: 159-228.

Iljin, B. S. 1927b. Opredelitel' bychkov (fam. Gobiidae) Azovskogo i Chernogo Morei. Trudy- Azov-Chernomorsk. Eksp., 2: pp. 128-143.

Iljin, B. S. 1956. Zamechaniya i popravki k podotryadu Gobioidei v knige L. S. Berga 'Ryby presnykh vod SSSR i sopredel'nykh stran', Izd. 4, 1948-1949, str. 1055-1125. Vop. Ikhtiol., 7: pp. 185-192.

Moreau, E. 1892. Manuel d'Ichthyologie française, Paris: viii+650 pp., III pl.

Nardo, G. D. 1847b. Sinonimia moderna delle specie registrate nell'opera inedita dell'Ab. Stefano Chiereghin intitolata: 'Descrizione de' crostacei, de' testacei e de' pesci che abitano le lagune e golfo veneto rappresentati in figure a chiaro-scuro e colori'. Venezia.

Nardo, G. D. 1860. Prospetti sistematici degli animali delle provincie Venete et del mare Adriatico, e distinzione delle specie in gruppi relative alla loro geografia fisica. Venezia, 104 pp.

Ninni, A. P. 1938. I Gobius del mari e delle acque interne d'Italia. Memorie R. Com. talassogr. ital., (242): 169 p.

Otto, A. W. 1821a. Conspectus animalium quorundam maritimorum nondum editorum. Inaug. Dissert. Vratislaviae, 20 p.

Otto, A. W. 1821b. Animalium quorundam maritimorum nondum editorum. Pars prior. Isis (Oken), 2, Heft 9, Beylage 18: 5 p. n. num.

Pallas, P. S. 1811 [probably 1814]. Zoographia rosso-asiatica, sistens omnium animalium in extenso Imperio Rossico et adjacentibus maribus observatorum recensionem, domicilia, mores et descriptiones anatomen atque icones plurimorum, 3 vol. Petropoli, 3, 428 pp. (pl. [1], XIII, XIV, XV, XXI, XX (sic)) (89 pl. are mentioned in the text but are unpublished. Perhaps published in Icones ad Zoographiam Rosso-Asiaticam, 1834-427.). (2nd ed., 1831, 3 vol. and Atlas, Petropoli.)

Risso, A. 1826. Histoire naturelle des principales productions de l'Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes. Paris et Strasbourg, vol. III, XVI+486 pp., 16 pl.

Smitt, F. A. 1899. Preliminary notes on the arrangement of the genus Gobius, with an enumeration of its European species. Öfvers. svenska VetenskAkad. Handl., 1899, 56 (6): pp. 543-555.

Soljan, T. 1963. Fishes of the Adriatic (Ribe Jadrana). Fauna et flora adriatica 1 (revised and enlarged for the English edition), 428 pp., many unnumbered fig.

Svetovidov, A. N. 1964. Rîbî Chernogo Morya. [The fishes of the Black Sea]. Opred Faune SSSR, 86: pp. 1-552, fig. 1-191 (in Russian).

Whitley, G. P. 1935. Studies in ichthyology. No. 9. Rec. Aust. Mus., 19 (4): pp. 215-250, 11 fig., pl. 18.

Zambriborsch, F. S. 1968. K sistematike bychkov Chernogo i Azovskogo Morei. Vest. Zool. Ak. Nauk. Ukr. S.S.R., 1968, (1): pp. 37-44.

Zei, M. 1963. Vertebrata (Cyclostomata, Chondrichthyes, Osteichthyes): 496-565, pl. 176-215, in R. Riedl, Fauna und Flora der Adria, Hamburg.

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