(H. Milne Edwards, 1834)

The carapace is slightly convex dorsally and flat medially; it is covered with a mosaic of tubercles, closely confluent, petaloid on the margins and with the centre slightly raised; all have blunt aspects; the furrows separating the regions are narrow and feebly demarcated. Attains 19.7 mm carapace breadth (1.3 times broader than long). Male chelipeds with black colouration of fixed finger not extending backwards onto palm. (Davie 1997)

Type locality: unknown.
Range: Red Sea (A. Milne Edwards, 1865, Odhner, 1925, Guinot, 1976, Serène, 1984); Suez (Klunzinger, 1913, Guinot, 1976, Serène, 1984); Gulf of Aden - Djibouti (Odhner, 1925); Mozambique - Delagoa Bay (Barnard, 1950); South Africa - Umhloti River mouth (Stebbing, 1905), off Umhloti River, Umtwalumi River (Barnard, 1950); Mozambique (A. Milne Edwards, 1865, Hilgendorf, 1879, Odhner, 1925); Madagascar - Nosy Faly, Majunga, Tany Kely and north-west coast (Serène, 1984); Seychelles (Rathbun, 1911); Mauritius (A. Milne Edwards, 1865, Michel, 1964); Saya de Malha Bank (Rathbun, 1911); Cargados Carajos Islands (Rathbun, 1911); Persian Gulf (Alcock, 1898); Pakistan - Karachi (Alcock, 1898); India - Tuticorin, Rameswaram and Madras (Henderson, 1893), Ganjam Coast (Alcock, 1898), Okha (Chhapgar, 1957); Sri Lanka (Alcock, 1898); Mergui Archipelago (Alcock, 1898); Strait of Malacca (Alcock, 1898); Taiwan - Taiwan Strait (Odhner, 1925), Tingch'ieting (Lin, 1949); China - Hong Kong (Alcock, 1898, Odhner, 1925), Pingtan, Fujian Province (Shen, 1940b), Guangdong, Fujian and Zhejiang (Dai & Yang, 1991); Macclesfield Bank (Odhner, 1925); Sulu Archipelago - Jolo (Odhner, 1925); north Borneo (Odhner, 1925); Indonesia - Ternate (de Man, 1902a), Kepulauan Kai (Odhner, 1925); Torres Strait (Miers, 1886); Australia - (Alcock, 1898), Swan River, W. A. (White, 1848), Cape Jaubert (Rathbun, 1924c), Shark Bay, W. Australia (Odhner, 1925), off Cape York (Miers, 1886), Port Denison, Queensland (Haswell, 1882c), Port Stephens (Haswell, 1882c), Port Jackson, N.S.W. (A. Milne Edwards, 1865, Haswell, 1882c), Moreton Bay (Campbell & Stephenson, 1970); New Caledonia (A. Milne Edwards, 1873, Odhner, 1925); between high and low tidal marks.

Kingdom Animalia
Phylum Arthropoda
Class Crustacea
Order Decapoda
Suborder Pleocyemata
Infraorder Heterotremata
Superfamily Xanthoidea
Family Xanthidae
Subfamily Actaeinae
Genus Actaea
Species Actaea savignyii

Status in World Register of Marine Species

Not included in WoRMS database

Scientific synonyms and common names

Cancer granulatus (not Linnaeus, 1758) Audouin, 1826: 87.
Cancer savignyii Milne Edwards H., 1834: 378.
Actaea granulata (part = savignyii) Milne Edwards A., 1865c: 275(part). -- Milne Edwards A., 1873a: 192. -- Miers, 1879b: 30. -- Haswell, 1882c: 44. -- Miers, 1886: 120. -- Henderson, 1893: 356. -- Alcock, 1898: 151. -- de Man, 1902a: 617. -- Stebbing, 1905: 30.
Euxanthodes granulata Paulson, 1875: 42, pl. 6, fig. 3.
Actaea Savignii Kossmann, 1877: 25, pl. 3, fig. 8. -- Hilgendorf, 1879: 787.
Actaea calculosa (not H. Milne Edwards, 1834) Nobili, 1906b: 261(part).
Actaea savignyi Rathbun, 1911: 221. -- Balss, 1924a: 8(part). -- Rathbun, 1924c: 17. -- Odhner, 1925: 52(part). -- Fox, 1927: 218(part). -- Monod, 1937: 18. -- Monod, 1938: 129. -- Shen, 1940b: 72, 85. -- Lin, 1949: 22. -- ? Barnard, 1950: 231, fig. 43b, c. -- Tortonese, 1952: 4(part). -- Holthuis, 1956: 326(part). -- Chhapgar, 1957b: 33, pl. 9 p-r. -- Michel, 1964: 21. -- Guinot, 1976: 211, figs 38A, 39A, 39a, 41A, pl. 9, figs 1-3. -- Serène, 1984: 109(key), 111(key), 113, fig. 66, pl. 14, fig. B. -- Dai & Yang, 1991: 308, pl. 39(7).
Actaea (Euxanthodes) Savignyi Klunzinger, 1913: 178, 194, pl. 5, fig. 6, pl. 7, fig. 4, pl. 10, fig. 8.
Actaea savignii Campbell & Stephenson, 1970: 278, fig. 37.

Non Actaea granulata -- Ortmann, 1893b: 455.
= Actaea semblatae Guinot, 1976.

Non Actaea granulata -- Nobili, 1906c: 127.
= Actaea jacquelinae Guinot, 1976.

Non Actaea savignyi -- Sakai, 1939: 485, fig. 37, pl. 61, fig. 2. -- Sakai, 1954: 75. -- Sakai, 1965b: 145, pl. 72, fig. 2. -- Takeda & Miyake, 1968c: 555. -- Kim, 1973: 383, 631, fig. 147, pl. 83, fig. 111a-b. -- Sakai, 1976a: 442, pl. 158, fig. 2. -- Yamaguchi et al., 1976: 38.
= Actaea semblatae Guinot, 1976.

Non Actaea savignyi -- Stephensen, 1945: 151, fig. 38C.
= ? Actaea jacquelinae Guinot, 1976.

Savigny-ohgigani [Japanese]
ohgigani, Savigny- [Japanese]

Alcock, A.W., 1898. Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part 1. The Family Xanthidae. Journal of the Asiatic Society of Bengal, Calcutta, 67, part 2(1): 67-233.

Audouin, J.V., 1826. Explication sommaire des Planches de Crustacés de l'Égypte et de la Syrie, publiées par Jules-César Savigny, membre de l'Institut; offrant un exposé des charactères naturels des genres, avec la distinction des espèces. In: J.-C. Savigny, Description de l'Égypte ou receuil des observation et des recherches qui ont été faites en Egypte pendant l'expedition de l'armée française 1798-1801. Histoire naturelle, 1(pt. 4): 77-98.

Balss, H., 1924a. Decapoden des Roten Meeres. III. Die Parthenopiden, Cyclo- und Catometopen. In: Expedition S.M. Schiff "Pola" in das Rote Meer, nördliche und südliche Hälfte 1895/96-1897/98. Zoologische Ergebnisse. XXXIV. Denkschriften der kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-naturwissenschaftliche Klasse, 99(6): 1-18, fig. 1.

Barnard, K.H., 1950. Descriptive catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum, 38: 1-837, figs 1-154.

Campbell, B.M. & W. Stephenson, 1970. The sublittoral Brachyura (Crustacea: Decapoda) of Moreton Bay. Memoirs of the Queensland Museum, 15(4): 235-302, figs 1-49, pl. 22.

Chhapgar, B.F., 1957b. Marine Crabs of Bombay State. Taraporevala Marine Biological Station, Contribution number 1: v + 89 pp., pls A, B, 1-16.

Dai, A. & S. Yang, 1991. Crabs of the China Seas, i-iv, 1-608, figs 1-295, pls 1-74. China Ocean Press, Beijing and Springer-Verlag, Berlin Heidelberg New York Tokyo, English edition. (Translation from Chinese original 1986.)

de Man, J.G., 1902a. Die von Herrn Professor Kükenthal im indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kükenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo, in Aufträge der Senckenberg. Naturforsch. Gesellschaft ausgeführt von Dr. Willy Kükenthal. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25(3): 465-929, pls 19-27.

Fox, H.M., 1927. Appendix to the Report on the Crustacea Decapoda (Brachyura). Zoological Results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22: 217-219.

Guinot, D., 1976. Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La superfamille des Bellioidea Dana et trois sous-familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mémoires du Muséum national d'Histoire naturelle, Paris, séries A, 97: 1-308, figs 1-47, pls 1-19. (in French)

Haswell, W.A., 1882c. Catalogue of the Australian stalk- and sessile-eyed Crustacea. The Australian Museum, Sydney: i-xxiv, 1-324 + 2 p. addenda, figs 1-8, pls 1-4.

Henderson, J.R., 1893. A contribution to Indian Carcinology. The Transactions of the Linnean Society of London, (series 2, Zoology) 5(1): 325-458, pls 36-40.

Hilgendorf, F., 1879. Die von Herrn W. Peters in Moçambique gesammelten Crustaceen. Monatsbericht der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1878: 782-851, pls 1-4.

Holthuis, L.B., 1956. Notes on a collection of Crustacea Decapoda from the Great Bitter Lake, Egypt, with a list of the species of Decapoda known from the Suez Canal. Zoologische Mededelingen, Leiden, 34(22): 301-330, figs 1-3.

Kim, H.S., 1973. A Catalogue of Anomura and Brachyura from Korea. In: Illustrated Encyclopedia of Fauna and Flora of Korea, Samhwa Publishing Company, 14: 1-694, figs 1-265, pls 1-112, tabls 1-2, 1 map. Seoul. (In Korean with English summary)

Klunzinger, C.B., 1913. Die Rundkrabben (Cyclometopa) des Roten Meeres. Abhandlungen der kaiserlich Leop.-Carol. Deutschen Akademie der Naturforscher Halle, 99(2): 97-402, figs 1-4, pls 5-11.

Kossmann, R., 1877. Zoologische Ergebnisse einer Reise in die küstengebiete des Rothen Meeres. Malacostraca. 1. Theil: Brachyura. Leipzig, W. Engelmann: 1-66, pls 1-3.

Lin, C.C., 1949. A catalogue of Brachyurous Crustacea of Taiwan. Quarterly Journal of the Taiwan Museum, 2(1): 10-33.

Michel, C., 1964. Checklist of the Crustacea Brachyura (Crabs) recorded from Mauritius. Mauritius Institute Bulletin, 6(1): 1-48.

Miers, E.J., 1879b. On a collection of Crustacea made by Capt. H.C. St. John R.N. in the Corean and Japanese Seas. Part I. Podophthalmia. With an Appendix by Capt. H.C. St. John. Proceedings of the Zoological Society of London, 1879: 18-61, pls 1-3.

Miers, E.J., 1886. Report on the Brachyura collected by H.M.S. Challenger during the years 1873-1876. In: C.W. Thompson & J. Murray, Report on the Scientific Results of the exploring Voyage of H.M.S. Challenger during the years 1873-1876, under the command of Captain George S. Nares, R.N., F.R.S. and the Late Captain Frank Tourle Thomson, R.N., Zoology, 17(2): i-xli, 1-362, pls 1-29.

Milne Edwards, A., 1865c. Études zoologiques sur les Crustacés récents de la famille des Cancériens. Cancérides, Pirimélides, Carpilides, Première Partie. Nouvelles Archives du Muséum national d'Histoire naturelle, Paris, 1: 177-308, pls 11-19.

Milne Edwards, A., 1873a. Recherches sur la faune carcinologique de la Nouvelle-Calédonie. Part II. Groupe des Cyclométopes Portuniens. Nouvelles archives du Muséum national d'Histoire naturelle, Paris, 9: 155-332, pls 4-18.

Milne-Edwards, H., 1834. Histoire Naturelle des Crustacés, comprenant l'anatomie, la physiologie et la classification de ces animaux. Libraire Encyclopédique de Roret, Paris, Vol. 1: i-xxxv, 1-468, pls 3, 5-6, 15-17, 20, 22-23, 25. (See Holthuis, 1979, for dates of publication.)

Monod, T., 1937. Crustacés. In: Mission A. Gruvel dans le Canal de Suez. I. Mémoires de l'Institut d'Egypte, 34: 1-19, figs 1-11.

Monod, T., 1938. Decapoda Brachyura. In: Mission Robert Ph. Dollfus en Egypte. VIII. Mémoires présentés a l'Institut d'Egypte, 37: 91-162, figs 1-29.

Nobili, M.G., 1906b. Faune carcinologique de la Mer Rouge. Décapodes et Stomatopodes. Annales des Sciences naturelles, Zoologie, (9) 4(1-3): 1-347, figs 1-12, pls 1-11.

Nobili, M.G., 1906c. Mission J. Bonnier et Ch. Pérez (Golfe Persique 1901). Crustacés Décapodes et Stomatopodes. Bulletin scientifique de la France et de la Belgique, 40: 13-159, figs 1-3, pls 2-7.

Odhner, T., 1925. Monographierte Gattungen der Krabben-familie Xanthidae. I. Göteborgs Kungliga Vetenskaps- och Vitterhets-Samhälles Handlingar, (4) 29(1): 1-92, figs 1-7, pls 1-5. (in German)

Ortmann, A.E., 1893b. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berücksichtigung der von Herrn Dr. Döderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur zeit im Strassburger Museum aufbewahrten Formen. Theil VII. Abteilung Brachyura (Brachyura genuina Boas), II. Unterabteilung: Cancroidea, 2. Section: Cancrinea, 1. Gruppe: Cyclometopa. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 7: 411-495, pl. 17.

Paulson, O.M., 1875. Izledovaniya rakoobraznykh krasnago morya s zametkami otnositel 'no rakoobraznykh drugikh morei. Tchasst I. Podophthalmata i Edriophthalmata (Cumacea). (Studies on Crustacea of the Red Sea with notes regarding other Seas, Part I). Kiev Kul'zhenko, i-xiv, 1-144, pls 1-21. (In Russian)

Rathbun, M.J., 1911. Marine Brachyura. In: The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the Leadership of Mr J. Stanley Gardiner. Vol. III, part II, No. XI. Transactions of the Linnean Society of London, (Zool.), (2) 14(2): 191-261, figs 1-2, pls 15-20.

Rathbun, M.J., 1924c. Brachyura, Albuneidae and Porcellanidae. In: Results of Dr. E. Mjöberg's Swedish Scientific Expeditions to Australia 1910-1913. 37. Arkiv för Zoologi, 16(23): 1-33, figs 1-7, pl. 1.

Sakai, T., 1939. Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha, pp. 365-741, figs 1-129, pls 42-111, table 1. Yokendo Co., Tokyo.

Sakai, T., 1954. Preliminary report on the Brachyuran Crustacea of Hachijo Island. Records of Oceanographic Works in Japan, 1(2): 73-77.

Sakai, T., 1965b. The Crabs of Sagami Bay, collected by His Majesty the Emperor of Japan, i-xvi, 1-206 (English text), figs 1-27, pls 1-100: 1-92 (Japanese text): 1-26 (references and index in English): 27-32 (index in Japanese), 1 map. Maruzen Co., Tokyo.

Sakai, T., 1976a. Crabs of Japan and the Adjacent Seas. (In 3 volumes: (1) English text: i-xxix, 1-773, figs 1-379, (2) Plates volume: 1-16, pls 1-251, (3) Japanese text: 1-461, figs 1-2, 3 maps.) Kodansha Ltd, Tokyo.

Serène, R., 1984. Crustacés Décapodes Brachyoures de l'Océan Indien occidental et de la Mer Rouge, Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par Crosnier (A): Carpiliidae et Menippidae. Faune Tropicale, no. XXIV: 1-349, figs A-C + 1-243, pls 1-48. (In French) (Translated into English by R.W. Ingle).

Shen, C.J., 1940b. On the collection of Crabs of South China. Bulletin of the Fan Memorial Institute of Biology (Zool.), Peiping, 10(2): 69-104.

Stebbing, T.R.R., 1905. South African Crustacea. Part III. In: Marine Investigations in South Africa, Cape Town, Cape Times Ltd, 4: 21-123, pls 17-26.

Stephensen, K., 1945. The Brachyura of the Iranian Gulf with an Appendix: The male pleopod of the Brachyura. In: Danish Scientific Investigations in Iran, Part 4. Copenhagen, E. Munksgaard: 57-237, figs 1-60.

Takeda, M. & S. Miyake, 1968c. Crabs from the East China Sea. I. Corystoidea and Brachygnatha Brachyrhyncha. Journal of the Faculty of Agriculture, Kyushu University, 14(4): 541-582, figs 1-11, pl. 6.

Tortonese, E., 1952. Some field-notes on the fauna of the Suez Canal (Timsah and Bitter Lakes). Ist. Univ. Fen. Fak, hidrobiol., ser. B, 1(1): 1-6.

Yamaguchi, T., M. Takeda & K. Tokudome, 1976. A list of crabs collected in the vicinity of the Aitsu Marine Biological Station and a preliminary report on the cheliped asymmetry of the crabs. Calanus, 5: 31-46, figs., pl., 2 tabs.

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