Tan & Ng, 1995
HOLOTYPE MALE. Carapace 1.4 times broader than long, regions not well-defined; dorsal surface covered with carapace 1.4 times broader than long, regions not well defined; dorsal surface covered with large pearly, mushroom-like granules which may be coalesced; groove parallel to carapace border broad; rim around border of carapace with scattered shallow granule-lined circular depressions, short rows of granules extending from rim towards centre of carapace; distinct, narrow median longitudinal ridge running posteriorly to half carapace length; 2 short, oblique row of granules immediately behind posterior end of ridge, anterior to branchial humps; branchial regions raised and covered with irregularly shaped granulated grooves amongst large mushroom-shaped granules; deep granule-lined groove on either side of cardiac and intestinal regions. Front slightly produced and upturned, moderately bibbed, with longitudinal depression immediately after frontal cleft; postfrontal prominences triangular with apices pointing towards centre of carapace; lateral borders of carapace with 3 shallow clefts, separating borders into 4 broad lobes; margins of hepatic and sub-hepatic regions protruded to form rounded angles; shallow, triangular groove on ventral surface, posterior to sub-hepatic angle; branchiostegite with distinct patch of well-spaced mushroom-like granules; postero-lateral border beside cardiac region protruded to form lobe. Basal segment of antennule covered with small granules and entirely occupies fossa when closed. Third maxillipeds densely covered with granules; merus with narrow and pointed apex, outer edge slightly convex, inner edge lined by large granules; ischium 1.9 times longer than merus along inner margin, inner edge lined by large granules, with larger granules proximally. Surface of chelipeds covered with small raised granules; outer surface of chelipeds more granular than inner surface; fingers short, palm 1.2 times longer than fingers; immovable finger with transverse parallel rows of small granules on outer surface forming shallow grooves in between, inner surface slightly concave along cutting edge; inner border of outer surface of palm with larger granules than rest of palm; edges of carpus lined by scattered large granules; large granules present close to proximal end of merus; cutting edges of both fingers sparsely lined by small, denticulate teeth. Outer surface of last pair of legs coarsely granulated, inner surface with flatter granules; entire posterior edge of merus lined with row of large granules; dactylus short and stout, covered with small granules. Abdomen densely covered with large pearly granules; segment 6 very broad, lateral margins expanded beyond general outline of abdomen; terminal segment with rounded apex. G1 stout, 2.3 times longer than G2, distal half slightly bulbous and sparsely covered with short hairs, apex simple with triangular opening; G2 narrow, with long and thin petaloid terminal process. (Tan & Ng, 1995)
Type locality: Kii Peninsula, Shionaomisaki, Japan.
Range: Japan (Takeda, 1979a).