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(H. Milne Edwards, 1834)

The antero-lateral margins of the carapace have 4 to 5 teeth behind the exorbital angle. The carapace and chelipeds have their surfaces granular or smooth, sometimes finely punctate. There are 5 teeth behind the exorbital angle; the fifth is sometimes reduced to a granule. The posterior antero-lateral tooth is much smaller that the others but quite distinct. The first pleopod of the male has 3-4 feeble, straight, subdistal spines and a short apical lobe bordered on one side by 5-6 mushroom-shaped tubercles for most of its length and followed distally by 2-3 curved spines. (Serène, 1984)

Type locality: Mauritius.
Range: Red Sea; Zanzibar (Lenz, 1905); Madagascar - Tuléar and Fort-Dauphin (Serène, 1984); Seychelles - Praslin and Coëtivy Islands (Rathbun, 1911); Mauritius (H. Milne Edwards, 1834, Bouvier, 1915b, Michel, 1964); Persian Gulf (Alcock, 1898); Laccadive Islands (Alcock, 1898); Sri Lanka (Alcock, 1898); Chagos Archipelago - Salomon and Peros, Coin (Rathbun, 1911); Andaman Islands (Alcock, 1898); Nicobar Islands (Alcock, 1898); Sumatra - Banda Aceh; Japan - Okinawa (Stimpson, 1907), Miyazaki, Okinawa and Northern Daito-jima (Sakai, 1939), Hachijo-jima (Sakai, 1954), Sagami Bay (Sakai, 1965b), Sagami Bay, Kii Minabe, Amami-Oshima, Yoron, Yaeyama-retto and Ogasawara-shoto (Sakai, 1976a), Kushimoto (Miyake, 1983), Hachijo-jima, Nanki Shirahama, Takara-jima, Yoron-jima, and Ishigaki-jima (Muraoka, 1998); Taiwan - (Parisi, 1916), Kaohsiung, Nanfangao, Chi-lung, and Su-ao (Lin, 1949); China - Paracel Island (Chen & Lan, 1978), Hainan Island and Paracel Islands (Dai & Yang, 1991); Philippines - Zamboanga (Miers, 1886); Indonesia - west Celebes; Palau (Takeda & Hayashi, 1973, Takeda, 1976a); New Ireland and Nissan Island (Sendler, 1923); Vanuatu - Torres Islands and Santo (Takeda & Nunomura, 1976); Fiji (Dana, 1852); Hawaiian Islands - Sandwich Island and Hilo (Dana, 1852), Oahu, Hawaii, Maui, and Necker Island (Rathbun, 1906), Sandwich Island (Stimpson, 1907); Line Islands - Palmyra and Tabuaeran (Edmondson, 1923); Tahiti (Sendler, 1923); Tuamotu Archipelago - Manihi (Dana, 1852), Makatea (Sendler, 1923).

Leptodius sanguineus