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(H. Milne Edwards, 1834)

The antero-lateral margins of the carapace have 4 to 5 teeth behind the exorbital angle. The carapace and chelipeds have their surfaces granular or smooth, sometimes finely punctate. There are 4 teeth behind the exorbital angle. The carapace has projecting regions that are rugose and separated by deep grooves. The first pleopod of the male has six stout, short subdistal spines and a noticeably long apical lobe with 8-10 mushroom-shaped marginal tubercles. (Serène, 1984)

Type locality: unknown.
Range: Red Sea (Kossmann, 1877, Klunzinger, 1913, Guinot, 1964b); Gulf of Aden - Djibouti (Guinot, 1964b); Persian Gulf (Alcock, 1898, Stephensen, 1945); Zanzibar (A. Milne Edwards, 1868); Mozambique - Delagoa Bay (Barnard, 1950); South Africa - Natal (Krauss, 1843); Moyotte (Guinot, 1958); Madagascar - Nosy Bé (Guinot, 1968a), Nosy Bé, Tuléar, Tamatave and Fort-Dauphin (Serène, 1984); Seychelles - (Rathbun, 1911), Aldabra Island (Guinot, 1964b, 1968a); Saya de Malha Bank (Rathbun, 1911); Mauritius (Richters, 1880, Bouvier, 1915b, Michel, 1964); Pakistan - Karachi (Alcock, 1898); India - Bombay (Alcock, 1898, Chhapgar, 1957); Sri Lanka (Alcock, 1898); Andaman Islands (Alcock, 1898); Burma - Akyab (Alcock, 1898); Mergui Archipelago (Alcock, 1898, Chopra & Das, 1937); Malaysia - Pinang (Alcock, 1898, Lanchester, 1902); Japan - Tokyo Bay and Sagami Bay (Ortmann, 1893), Hondo and Goto-retto (Parisi, 1916), Tanabe, between Nagasaki and Koshiki-jima, and north-west of Nagato (Yokoya, 1933), Shimoda (Sakai, 1935), Tateyama Bay, Misaki, Shimoda, Nagasaki and Okinawa (Sakai, 1939), Hachijo-jima (Sakai, 1954), Sagami Bay (Sakai, 1965b), from Boso Province to Okinawa and Ogasawara-shoto (Sakai, 1976a), Amakusa (Yamaguchi et al., 1976), Yaeyama-retto (Miyake, 1983), Okukubi River, Okinawa (Shokita et al., 1998), Ginowan, Okinawa (Shokita et al., 2000); Korea - Cheju-do (Kamita, 1941), Cheju-do and Chuja Island (Kim, 1973); Taiwan (Lin, 1949); China - Qingdao (Shen, 1936), Xiamen (Shen, 1940b), Guangdong, Fujian, and Paracel Islands (Dai & Yang, 1991); Philippines - Zamboanga and Negros (Garth & Kim, 1983); Indonesia - Noordwachter Island (de Man, 1887d), Ambon (Rathbun, 1910b), Pontianak; Palau (Sendler, 1923, Takeda, 1976a); Vanuatu - Santo (Takeda & Nunomura, 1976); New Caledonia - Nouméa, Ilot Maitre, Plum, and Hienghéne (Takeda & Nunomura, 1976); Gilbert Islands (Holthuis, 1953); Hawaiian Islands - Honolulu, Pearl Harbor and Hilo (Rathbun, 1906); Tuamotu Archipelago (Holthuis, 1953); between high and low tidal marks.

Leptodius exaratus