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Williams, G.C., 1995. Living genera of sea pens (Coelenterata: Octocorallia: Pennatulacea): illustrated key and synopses. Zoological Journal of the Linnean Society 113: 93-140.

Williams, G.C., 1999. Index Pennatulacea Annotated Bibliography and Indexes of the Sea Pens (Coelenterata: Octocorallia) of the World 1469-1999. Proceedings of the California Academy of Sciences 51 (2): 19-103, 14 plates, 1 figure.

Genus Virgularia